How to complete the Toward the Nest puzzle in Crimson Desert

I spent a long time trying to solve the Toward the Nest puzzle in Crimson Desert—far more than any I'd encountered in the main story up to that point. By comparison, finishing Abyss Without Balance or activating the Spire of the Stars was super easy. In this quest, you'll find yourself chasing the Crowcaller into the Abyss, and surprise, surprise, there's a puzzle.

You'll first have to ascend the Spire of Soaring and complete some relatively easy circuit puzzles along the way. But where...